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Assuming we have a page, which has as Status Code 200 OK and as Response Headers Status HTTP/1.1 410 Gone, like on image:

enter image description here

What is the behavior of bots / Googlebot? To crawl/index or not?

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Bots should see the same as what you are seeing in the browser and interpret the response in the same way. ie. as a 200 OK. That is, after all, what the HTTP Status-Line (the first line of the response header) is reporting - which is part of the HTTP standard.

The Status HTTP response header is non-standard. (It's less common to see the "HTTP version" specified in this field as well?)

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